Dialogue Numbering can be locked, numbering new speeches as A-Speeches—while reordered speeches maintain their original numbers. You might want to do if you're using these numbers to relate to story boards or cells.
Under the Production menu, select Animation Locking, then Lock Dialogue Numbering.
This causes a number for each speech to be displayed onscreen, to the right of the Character Name in angle brackets (e.g., <6>). (This number will not print unless you select Dialogue on the Print menu and check Dialogue Numbered/Laid Out for Animation Recording Scripts, above.) It is displayed for reference, to show what's going to print in case you want to override a Dialogue Number.